

Triceratops and Torosaurus have long been considered the kings of the horned dinosaurs. But a new discovery traces the giants' family tree further back in time, when a newly discovered species appears to have reigned long before its more well-known descendants, making it the earliest known member of its family.
The new species, called Titanoceratops after the Greek myth of the Titans, rivalled Triceratops in size, with an estimated weight of nearly 15,000 pounds and a massive eight-foot-long skull.
Titanoceratops, which lived in the American southwest during the late Cretaceous period around 74 million years ago, is the earliest known triceratopsin, suggesting the group evolved its large size more than five million years earlier than previously thought, according to Nicholas Longrich, the palaeontologist at Yale University who made the discovery. The finding, which will appear in an upcoming issue of the journal Cretaceous Research, helps shed light on the poorly understood origins of these giant horned dinosaurs.
Longrich was searching through scientific papers when he came across a description of a partial skeleton of a dinosaur discovered in New Mexico in 1941. The skeleton went untouched until 1995, when it was finally prepared and identified incorrectly as Pentaceratops, a species common to the area. When the missing part of its frill - the signature feature of the horned dinosaurs - was reconstructed for display in the Oklahoma Museum of Natural History, it was modelled after Pentaceratops.
'When I looked at the skeleton more closely, I realised it was just too different from the other known Pentaceratops to be a member of the species,' Longrich said, adding that the specimen's size indicated that it likely weighed about twice as much as adult Pentaceratops. The new species is very similar to Triceratops, but with a thinner frill, longer nose and slightly bigger horns, Longrich said.
Instead, Longrich believes that Titanoceratops is the ancestor of both Triceratops and Torosaurus, and that the latter two split several millions years after Titanoceratops evolved. 'This skeleton is exactly what you would expect their ancestor to look like,' he said.
Titanoceratops was probably only around for about a million years, according to Longrich, while the triceratopsian family existed for a total of about 10 million years and roamed beyond the American southwest into other parts of the country and as far north as Canada.
In order to confirm the discovery beyond any trace of a doubt, Longrich hopes palaeontologists will find other fossil skeletons that include intact frills, which would help confirm the differences between Titanoceratops and Pentaceratops.
'There have got to be more of them out there,' Longrich said.
Study of oldest turtle fossil shows how shell evolved
Glacial erosion changes mountain responses to plate tectonics
Small juvenile dinosaur fossil sheds light on evolution of plant eaters
New bizarre feathered dinosaur discovered in China